The game starts out with a recap of Lords of Shadow and Mirror of Fate, detailing Gabriel Belmont's (Robert Carlyle) transformation into Dracula and his encounters with his son, Trevor (Richard Madden), whom he kills and subsequently resurrects, and grandson, Simon. The game starts proper with knights attacking Dracula's castle. He swiftly dispatches all of them before being confronted by the revived Trevor, now known as Alucard.
A thousand years have passed and Dracula awakens in a cathedral in
modern times, weak from starvation and with no memory of the
circumstances that led him there. He is warned by his former enemy,
Zobek (Patrick Stewart),
of the return of Satan, who until then was deterred from doing so by
Dracula himself. Zobek offers him his old weapon, the Vampire Killer,
the only relic capable of ending his cursed existence, in exchange for
him vanquishing the people trying to bring Satan (Jason Isaacs)
back, his acolytes. The key to him regaining his power lies in his
castle, which the cursed blood that kept it alive for centuries attempts
to deter Dracula of his quest. Throughout his journey, Dracula is aided
by various allies, including Zobek's bodyguard and specters of his
deceased wife Marie (Natascha McElhone) and a younger Trevor. He also encounters the Brotherhood of Light and its current leader, Victor Belmont (Anthony Howell).
Having destroyed two of the acolytes and tracked down the third,
Dracula prepares to ambush him but is stopped by the bodyguard,
flashbacking to his last encounter with Alucard. It is revealed that
centuries ago, Alucard had come to Dracula with a plan to destroy both
Zobek and Satan, using his sword, the Crissaegrim, on him to induce a
deep sleep and thus setting the current events into motion. The truth
now known, the bodyguard reveals himself to be Alucard. Zobek discovers
their treachery and angrily confronts them both. Dracula kills him just
as the third acolyte summons Satan, who proceeds to summon a flying
monster to destroy the human world. Dracula and Alucard are able to kill
the beast, only for Satan to possess the latter, forcing father and son
to battle. Dracula gains the upper hand and prepares to stake Alucard
with the Vampire Killer, despite Satan's insistence that he would not
and forcing him to vacate the body. Having anticipated this, Dracula
kills Satan once and for all while saving his son. Dracula then destroys
the Mirror of Fate and walks back inside his cathedral with Alucard as
the sun rises on the city.
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